Another rust

Walking through Hoe Wood, I noticed the Lords-And-Ladies Arum maculatum were dotted with pale orange spots.

Looking under the leaves, I spotted tell-tale signs of another rust fungus from the genus Puccinia. Forming circular pustules at first, then bursting out into a ring of cup-shaped aecia.

Ellis & Ellis (1997) lists only one Puccinia species on Arum: Puccinia sessilis.

For the record
Date: 11/04/2019
Location: Hoe Wood [private site]